Each Lemmy instance keeps a copy of posts and contents made in communities followed by its users, so even though an instance is decommissioned, the content lives on on other instance.
For example my instance was off for a month (I was moving out, setting up my NAS was my last priority) and during this time my posts and comments were still available on other instances.
As for users unable to login on decommissioned instances, unfortunately all they can do is create an account on another instance.